Short post. Map in HD at this URL: https://i.imgur.com/rqn7zs3.png

CONTENTS:

  1. Intro
  2. Background info
  3. Corded Ware sample info
  4. The admixture analysis
    4.1. Corded Ware
    4.2. Yamnaya
  5. WSH ancestry map
  6. Global25 coordinates

1. Intro

Earlier this year I published an admixture analysis showing the levels of Yamnaya-like ancestry throughout Eurasia. Unfortunately, the genetic model I used was missing key ancient populations, so the prediction was not accurate. I wasn’t familiar with the software so had to use someone else’s model. Since then, I’ve experimented a lot with Vahaduo and have created a model of my own using the Global25 calculator. I’ll include all of the G25 coordinates/samples at the end of the article.

2. Background info

Alongside Western Steppe Herders (Yamnaya and early Corded Ware), my model features the following ‘basal’ West and Central Eurasian populations of the early Holocene:

  • East European Hunter-Gatherers (EHG)
  • Scandinavian Hunter-Gatherers (SHG)
  • Baltic Hunter-Gatherers (SHG)
  • West European Hunter-Gatherers (WHG)
  • West Siberian Hunter-Gatherers (WSHG)
  • Anatolian Neolithic Farmers (ANF)
  • Caucasus Hunter-Gatherers (CHG)
  • Iranian Neolithic Farmers (Iran_N)
  • Natufians (Levantine Farmers)
  • Iberomaurusians (Ancient North Africans, from Taforalt)
  • East Asian Siberians (from Krasnoyarsk)

This gives a more accurate picture of Western Steppe Herder (WSH) ancestry in Europe than the classic three-way model (Yamnaya + EEF + WHG), because the three-way model often groups a lot of European hunter-gatherer ancestry with WSH, which throws off the figures.


European hunter-gatherers populations were differentiated by their levels of West European Hunter-Gatherer and Ancient North Eurasian ancestry, which varied throughout the WHG/EHG region; some populations had more ANE, others more WHG.

The Krasnoyarsk East Asian Siberian was near-identical to modern Samoyedic Nganasan people, while ‘West Siberian Hunter-Gatherer’ (WSHG) peoples were most similar to modern Besermyans, Udmurts, or Saamis (Europeans with ~20% East Asian Siberian ancestry). We have no idea what WSHG looked like, but it was probably something like this:

3. Corded Ware sample info

The Corded Ware sample (“PNL001”) featured in this admixture analysis represents the earliest type of Corded Ware population, from before they extensively intermixed with Early European Farmers and European Hunter-Gatherers. The sample was located in Bohemia and dates to around approximately 2900 BC.

The genetic distance between the Early Corded Ware and Yamnaya populations was similar to the distance between Germans and Icelanders, i.e. negligible. They are best described as sister populations or close cousins.

While the Yamnaya lived on the Eastern European steppe, Corded Ware likely originated in the forest steppe to their north. (More on Corded Ware origins in this article).

The PNL001 individual was 96% identical to Yamnaya, with only ~3% additional ancestry from a Baltic Hunter-Gatherer type population (from the WHG/EHG mixed zone) and ~1% from Anatolian Farmers. They also carried the Y-DNA haplogroup (male lineage) R1b-S21/U106, which is associated with modern Germanic peoples.

4. The admixture analysis

Please bear in mind that these are estimates and not 100% accurate.

4.1. Corded Ware

Here is my model when using Corded Ware as the WSH source population. Note that Northeastern Europeans, e.g. Finns, have increased hunter-gatherer ancestry that is not visible in the standard three-way model.

4.2. Yamnaya

Using Yamnaya as the WSH source population produces similar results, but the Corded Ware sample gives a better fit (lower ‘Distance’ number) for all populations.

Note: It is possible to further improve the overall fit of this model by using a wide variety of Late Neolithic and Bronze Age source populations that chronologically overlapped the Proto-Indo-European expansion. However, for ease of interpretation, I think it is best to stick to more basal populations.

5. WSH ancestry map

Here’s a map of the Corded Ware admixture analysis results:

6. Global25 coordinates

G25 source populations

WSH::Corded_Ware_CZE_early:PNL001.merged,0.12862,0.099522,0.050911,0.122741,-0.016926,0.046854,0.006345,-0.001846,-0.048677,-0.071619,-0.004709,0.001499,-0.008028,-0.014863,0.03393,0.008353,-0.01004,-0.003167,-0.006913,0.003126,-0.00262,0.002597,0.006532,0.02904,-0.00467
WSH::Yamnaya_RUS_Samara,0.1255849,0.089028,0.0426986,0.1153479,-0.0287232,0.0450564,0.0036033,-0.0025642,-0.0559032,-0.0728943,0.0018222,3.32e-05,-0.0026924,-0.0233041,0.0366141,0.0157633,-0.0012316,-0.0017879,-0.0038408,0.0137704,-0.0031749,0.0007557,0.0110649,0.0186102,-0.004537
EHG::RUS_Karelia_HG,0.1236877,0.0321583,0.129855,0.2101663,-0.010361,0.0571723,-0.0196627,-0.0234603,-0.002659,-0.0860153,0.0182957,-0.0184337,0.0333497,-0.039085,0.018865,0.0295237,-0.0148203,0.0031673,-0.0044413,0.012506,-0.007487,0.0169817,0.0093667,-0.021007,-0.0103383
SHG::SWE_Meso,0.1320345,0.0911438,0.1663102,0.19154,0.079938,0.0595432,-0.0013515,0.0118262,0.0377858,-0.0452402,0.0017052,-0.0192952,0.029435,-0.0175472,0.0356265,0.041368,-0.0016297,0.0007285,-0.0053738,0.0412698,0.0442342,0.0150548,-0.020459,-0.09176,0.0039815
BHG::Baltic_LVA_HG,0.1292603,0.1004104,0.1802636,0.1972329,0.1064236,0.0560919,0.0083574,0.0207106,0.0524476,-0.0261394,-0.0046788,-0.0181151,0.0281434,-0.0053759,0.0355759,0.0480141,0.003072,0.0040856,-0.0059236,0.042325,0.0613526,0.0130531,-0.0262979,-0.1110549,0.0095949
WHG::WHG,0.1246365,0.116278,0.184789,0.189279,0.1546445,0.0464355,0.0131605,0.0372675,0.0890705,0.017768,-0.0153455,-0.015811,0.0159065,-0.0030275,0.053338,0.0582065,0.00502,0.016343,-0.0093015,0.055589,0.0944585,0.0111905,-0.049607,-0.160866,0.0170045
ANF::TUR_Barcin_N,0.1175998,0.180118,0.0035312,-0.101158,0.0510443,-0.0483875,-0.0043582,-0.0069334,0.0362287,0.0807473,0.0079718,0.0118803,-0.0234545,0.0004691,-0.0419807,-0.0101913,0.0233091,0.0019866,0.0136954,-0.0097489,-0.0142249,0.0057723,-0.0041232,-0.0031658,-0.0043437
Iran_N::IRN_Ganj_Dareh_N,0.0430252,0.0664158,-0.1550722,0.0047158,-0.122669,0.0235384,0.017109,-0.0011998,-0.082546,-0.0544158,-0.0028258,-0.0016186,0.0044896,-0.0062756,0.0316498,0.0561384,-0.0054242,0.0068664,0.0136508,-0.0334162,0.00856,-0.028836,-0.0110678,-0.039331,0.0222254
CHG::GEO_CHG,0.091058,0.102568,-0.083344,-0.00323,-0.08617,0.020638,0.024911,-0.001846,-0.128236,-0.074717,-0.006333,0.023979,-0.054856,0.004404,0.026601,-0.03275,0.02386,-0.013429,-0.022249,0.034767,0.033815,-0.007048,0.006532,-0.025787,-0.002036
Natufian::Levant_Natufian,0.01935,0.135065,-0.039221,-0.135984,0.026774,-0.076137,-0.019036,-0.024691,0.100626,-0.008018,0.02858,-0.019633,0.067343,0.001651,0.022801,0.02612,-0.0103,0.006714,-0.018101,0.041395,-0.004118,-0.003215,-0.014297,-0.011206,0.011975
Iberomaurusian::MAR_Taforalt,-0.189857,0.0814452,-0.0242866,-0.085595,0.027636,-0.0552202,-0.0705968,0.0184146,0.155397,0.003499,0.0209156,-0.0318316,0.0747168,-0.0513334,0.0711988,-0.0363032,0.0052676,-0.066106,-0.1424162,0.0389938,-0.0376836,-0.1255322,0.0730118,-0.0137606,0.0164534
WSHG::RUS_Tyumen_HG,0.103579,-0.059916,0.100314,0.202199,-0.091094,0.044901,-0.048882,-0.054921,-0.035178,-0.090571,0.028093,-0.011989,0.02661,-0.077619,0.028637,0.01485,-0.015646,-0.004814,-0.004399,0.001376,-0.025705,0.011747,0.02465,0.00976,-0.013053
EastAsian_Siberian::RUS_Krasnoyarsk_BA,0.034147,-0.421445,0.150471,-0.002907,-0.14495,-0.082273,0.019741,0.035998,0.02577,0.000364,0.083792,-3e-04,0.015758,-0.048994,-0.04343,-0.029833,0.000652,0.005574,0.004902,-0.012381,0.026453,0.003462,0.001725,0.00723,0.012813

G25 target populations

Norwegian,0.1338234,0.1279567,0.0679357,0.0534334,0.0386443,0.0210366,0.0041964,0.0062964,0.0057267,0.0003906,-0.0037349,0.0036823,-0.007518,-0.011698,0.0210754,0.0093949,-0.0054203,0.0026061,0.0006823,0.0007503,0.005633,0.0055643,0.0042784,0.0177304,0.0010947
Swedish,0.1339317,0.1282953,0.0720659,0.0563097,0.0409013,0.0214348,0.0070612,0.0092964,0.0049085,-0.0065951,-0.0048022,0.001513,-0.004651,-0.0038337,0.0150003,0.0040156,-0.0072332,0.0020029,0.0034837,0.0048772,0.0070768,0.0022787,0.0011269,0.0106612,-0.0009751
Icelandic,0.1326987,0.1336267,0.067819,0.0563097,0.0393663,0.0229619,0.0033488,0.0085188,0.0026418,6.07e-05,-0.0054806,0.0051702,-0.0063675,-0.0090831,0.0230725,0.0052924,-0.0116909,0.0019109,0.0017389,0.0038873,0.0092545,0.0028028,-0.0008422,0.0140179,0.0008282
Scottish,0.1314657,0.1342312,0.0622654,0.0470426,0.039106,0.0172812,0.0035502,0.0051756,0.0035719,0.0028246,-0.005614,0.0050473,-0.0116806,-0.0119731,0.023024,0.0031254,-0.0112549,0.0027736,0.0025499,0.0014516,0.0042246,0.0033076,-0.0006559,0.0135561,-0.0007698
Irish,0.1333605,0.1340977,0.0611689,0.0488642,0.037788,0.0193517,0.0032928,0.0047129,0.0035683,0.0029265,-0.0069712,0.0058395,-0.0141892,-0.0140763,0.0259353,0.0052146,-0.0111456,0.0018944,0.0005974,0.0017758,0.0051145,0.0012787,0.0002929,0.0144074,0.0006607
Dutch,0.1267527,0.1313048,0.0610935,0.0461486,0.040469,0.0166244,0.0065435,0.0080802,0.003576,0.0003275,-0.0064447,0.0045873,-0.0087082,-0.0088422,0.0174443,0.005345,-0.0082143,0.0012669,0.0042698,0.0028138,0.0046909,0.0028672,0.0002678,0.0153541,-0.0004921
Danish,0.1305084,0.1354347,0.0661333,0.0517975,0.0413152,0.0192751,0.0056348,0.0078878,0.0030168,-0.0034914,-0.0054511,0.0039135,-0.0085547,-0.0060085,0.019112,0.0052132,-0.0072097,0.0020645,0.002931,0.0038881,0.0053683,0.0018154,0.0005826,0.0139805,0.000479
English,0.1318551,0.137043,0.0617883,0.044013,0.0392299,0.0167481,0.0049845,0.0057448,0.0052637,0.0056781,-0.0047691,0.0056161,-0.0125892,-0.0103507,0.0206153,0.0035521,-0.0103689,0.004094,0.003685,0.0029619,0.0059369,0.0034036,-0.0032693,0.0137938,3.16e-05
Welsh,0.1319779,0.1381628,0.0601319,0.0430236,0.0405922,0.0166218,0.0033135,0.006392,0.0062789,0.0043372,-0.0042302,0.0059122,-0.0129112,-0.0109754,0.0205954,0.005708,-0.0087032,0.0023564,0.003086,0.0020072,0.0049849,0.0038518,-1.24e-05,0.0112243,0.0008083
German,0.1302626,0.1373792,0.0579811,0.0387068,0.0400035,0.0156461,0.0040665,0.0057865,0.003526,0.00188,-0.0044133,0.0023732,-0.0053706,-0.0025417,0.0085125,0.0032913,-0.0032728,0.0016631,0.0033669,0.0017191,0.00314,0.002005,7.18e-05,0.0083052,0.0002774
Austrian,0.1290886,0.1350056,0.054394,0.032737,0.0347756,0.0125666,0.0072299,0.0062441,0.0028274,0.0027551,-0.0038781,0.0007847,-0.0012942,0.0027849,0.0005029,0.0036968,0.0002223,0.00079,0.0031793,0.0008385,-0.0009909,-0.0026548,0.0016675,0.0029627,-0.0010284
Belgian,0.1271867,0.1378863,0.0508694,0.0285197,0.038879,0.0112177,0.0042389,0.0034529,0.0074159,0.0110489,-0.0050521,0.0047014,-0.0127298,-0.0096131,0.0131448,0.0035701,-0.004404,0.00221,0.0022114,-0.0012784,0.0035491,0.0017494,-0.0014059,0.0102914,0.0011088

Estonian,0.1329454,0.1141455,0.087756,0.0833017,0.0427771,0.0297575,0.0113745,0.0144457,0.0004091,-0.028447,6.5e-05,-0.0123489,0.0194597,0.0213591,-0.0066639,0.0017104,0.0002216,-0.0024832,0.0013451,0.0022137,0.001385,-0.0018672,0.0042644,0.0007471,0.0030535
Latvian,0.1354494,0.122473,0.0887742,0.089471,0.043208,0.0340246,0.0125966,0.0140762,-0.002168,-0.0368842,-0.0035728,-0.0138178,0.0232208,0.031378,-0.010559,0.0040572,0.0027118,-0.0003802,0.0016088,0.0043022,-0.0027952,-0.0057128,0.0090708,-0.0093264,0.0026346
Lithuanian_VZ,0.1372991,0.1261795,0.0850879,0.082365,0.0444314,0.0326652,0.0104872,0.0176242,-0.0036048,-0.0315269,-0.0036131,-0.0097039,0.0199762,0.0301394,-0.0104505,-0.0045578,-0.0006029,0.0012828,0.0032525,0.0013445,-0.0043829,-0.0056728,0.00835,-0.005874,-0.001317
Belarusian,0.1322625,0.1236239,0.0750219,0.0695957,0.0402331,0.0265691,0.0107321,0.0133226,-0.0002727,-0.0246017,-0.0026957,-0.0092119,0.0193259,0.0272309,-0.0112647,-0.0038009,-0.0009647,0.0003717,0.0041397,-0.0010504,-0.0042342,-0.0046081,0.0077235,-0.0068601,0.0006946
Ukrainian,0.1309273,0.1240867,0.0683506,0.0575551,0.0376867,0.0212936,0.0087651,0.0113072,-0.002891,-0.0210999,-0.0016327,-0.0076594,0.0153563,0.0216328,-0.0112354,-0.0010965,0.0046903,-0.0006985,0.0025989,0.0017002,-0.0044989,-0.003479,0.0067387,-0.0040578,0.0007217
Polish,0.1318405,0.1292694,0.0698685,0.0577382,0.0406754,0.0217127,0.0086781,0.0108626,-0.0009329,-0.0185524,-0.0043488,-0.0064187,0.013093,0.0186295,-0.0070244,-0.0005595,0.0015455,-7.41e-05,0.0026948,0.0012872,-0.0031286,-0.0031185,0.0056063,-0.0033328,-8.76e-05
Czech,0.1311592,0.1318625,0.0604843,0.0472574,0.038658,0.0189217,0.0063451,0.0086623,0.0046724,-0.0088874,-0.0016862,-0.0038965,0.0060493,0.011772,0.0017435,0.0009995,-0.0006419,-0.0001462,0.0035679,0.0028956,-0.001267,-0.0001617,0.0065132,0.0039302,0.0015935
Hungarian,0.1270753,0.1296252,0.0553559,0.0364298,0.0363804,0.0119326,0.0066138,0.0075326,0.0020597,-0.0046731,-0.0014731,-0.0027191,0.0038544,0.0087488,-0.0043722,0.0001231,-8.4e-05,-0.0003891,0.0026486,-0.0013221,-0.003378,0.0013865,0.0055286,0.0026939,0.0004789
Ingrian,0.1297583,0.0900437,0.0898803,0.0756897,0.033032,0.0258437,0.009949,0.0129993,-0.0006137,-0.0265457,0.0031937,-0.0098913,0.0146183,0.0056883,-0.0029407,0.002696,-0.001608,-0.0005913,-0.0043573,0.004961,0.0040343,0.000165,0.007477,0.002008,0.0030337
Finnish,0.1305931,0.0918715,0.0899057,0.0766156,0.0317187,0.0232595,0.0087265,0.013184,0.0031769,-0.023071,0.0036809,-0.0060345,0.0119227,0.0058077,0.0021081,-0.0012815,-0.008136,0.0006589,0.0012905,0.0027597,0.0070458,0.0011706,0.0027937,0.0056955,0.0021076
Russian_Kostroma,0.1282407,0.0873355,0.0814582,0.0680453,0.027441,0.0215208,0.0119073,0.0129993,-0.0033747,-0.0299475,0.0042222,-0.0074932,0.017963,0.0123402,-0.0091383,-0.0039112,0.0009127,-0.0016258,-0.0017597,0.0004792,6.23e-05,-0.0014427,0.0035947,-0.0006025,-0.0020557
Karelian,0.1273094,0.0768108,0.093286,0.0808381,0.0275575,0.0224719,0.0102478,0.0152161,0.0017601,-0.0294117,0.0052308,-0.0110312,0.0189294,0.0089496,-0.00445,0.0013259,-0.0023746,-0.0014089,-0.0018435,0.0037822,0.0064659,-0.0009743,0.001154,0.0006975,0.0038138
Mordva_Erzya,0.127387,0.0819194,0.0728158,0.0672378,0.023 6452,0.0202195,0.0105755,0.0129611,-0.0045335,-0.0284745,0.0007984,-0.0105654,0.0176907,0.0121336,-0.0121582,2.21e-05,0.0043244,-0.0002008,0.0014664,0.0012297,-0.0025892,-0.0021846,0.0035125,-0.0064667,0.0015467
Vepsian,0.128051,0.0723847,0.0918707,0.0796554,0.0249447,0.0203745,0.0108626,0.0158712,0.0029428,-0.0290768,0.0050341,-0.0117979,0.0223569,0.0118126,-0.0086559,-0.0015395,-0.0013618,0.000183,-0.0014804,0.0035711,0.0042841,-0.0025281,0.0006231,0.0009839,0.0038651
Komi,0.1187174,0.0307705,0.0850408,0.0682822,-0.0046161,0.0141677,0.0094945,0.0130841,-0.0030884,-0.0298138,0.0099383,-0.0086022,0.0182258,-0.0062204,-0.0087676,-0.0047601,-0.0014733,-0.0014444,-0.0039595,-0.0039646,0.0014723,-0.0011993,-0.0001234,0.0012051,0.0004191
Udmurt,0.1067802188,-0.02780015625,0.07958421875,0.06548825,-0.03473715625,0.00783503125,0.0092828125,0.010925,-0.01241203125,-0.03636753125,0.0218311875,-0.00570428125,0.01547003125,-0.0341604375,-0.01067528125,-0.00502590625,-0.00275025,-0.00111646875,-0.01067646875,-0.0099656875,0.000662875,0.00326521875,-0.00467178125,0.0092745,-0.00298990625
Saami,0.1112621,-0.0302122,0.111722,0.079781,-0.0093286,0.008297,0.0081371,0.0138599,0.0029656,-0.0320964,0.0222877,-0.0074746,0.0180716,-0.0198263,-0.0040038,0.0026933,0.0011571,-0.0014886,-0.0059862,0.0044944,0.0158158,0.0013447,-0.0025649,0.0026584,0.0003442

Italian_Calabria,0.1028203,0.1462363,-0.011942,-0.0466197,0.0125147,-0.0117133,-0.0024283,-0.0031537,0.0044993,0.0215037,0.000433,0.0027977,-0.0055003,-0.000963,-0.009636,-0.0052593,0.0068237,0.0009713,0.0051537,-0.003043,-0.0026203,-0.0023907,-0.0001643,0.0037753,0.00012
Sicilian_West,0.1001643,0.1455593,-0.0003773,-0.03553,0.019388,-0.0098543,-0.0006267,0.0005383,0.013362,0.0222937,0.00341,0.0030973,-0.00223,0.0018807,-0.002262,-0.003182,-0.0018687,-4.23e-05,-0.00088,-0.006128,0.000832,0.0008243,-0.0032043,-0.0034943,0.0002393
Slovakian,0.1304414,0.128769,0.0626776,0.0514862,0.0412384,0.0200244,0.006768,0.0077998,-0.0018408,-0.0115538,-0.0019162,-0.0037466,0.009871,0.0104318,-0.0091476,0.0092814,0.012934,0.0002534,0.0054304,0.0023512,-0.0037186,-0.0020774,0.0043382,0.001928,-0.0018922
Slovenian,0.1288476,0.1352686,0.056191,0.0338504,0.0349602,0.0126616,0.0065332,0.0069688,0.0031086,-0.004483,-0.0031828,-0.0022482,0.0022298,0.0151936,-0.0086318,-0.002201,0.0002086,0.0013682,0.0078184,-0.0011508,-0.004018,-0.0027946,0.0014052,-0.001976,0.0021792
Croatian,0.128051,0.1338467,0.0518917,0.0314925,0.035822,0.0125223,0.0060161,0.0067381,0.0018408,-0.0058862,-0.001153,-0.0023827,0.0047275,0.0127851,-0.0066365,-0.0022011,0.0027512,-0.0007223,0.0018101,-0.0007503,-0.005166,-0.0024236,0.0044246,-0.0010966,0.0001796
Bosnian,0.1290754,0.1348622,0.0449528,0.0248064,0.0312058,0.0098726,0.004841,0.0052152,-0.0024136,-0.001859,0.0004224,-0.0028176,0.00556,0.0162396,-0.0125404,0.0010078,0.0066756,-0.000583,0.004827,0.0030266,-0.0050912,-0.0034376,0.0075182,0.001229,-0.0016286
Serbian,0.1273334,0.137229,0.040024,0.0148299,0.0314974,0.0045957,0.0048738,0.006672,-0.0011737,-7.9e-06,-0.0022946,-0.001251,0.0034128,0.012434,-0.0128875,-0.0008358,0.0040476,0.0005894,0.0043011,-0.0024468,-0.0083168,-0.0017151,0.0048925,0.000372,-0.0020097
Moldovan,0.1239316,0.1276665,0.0401454,0.0200645,0.0276461,0.0083535,0.0071733,0.0066151,-0.0009739,-0.0020306,-0.0017438,-0.0019697,0.0050333,0.0140867,-0.0108867,-0.0037,0.0041195,-0.0003559,0.0034149,-0.0011285,-0.0040672,0.0002709,0.0035095,-0.0004848,-0.0003193
Romanian,0.1245225,0.1340499,0.034167,0.0087856,0.0288362,0.0030677,0.0036425,0.0059073,0.0012475,0.0061596,-0.0034427,-0.0002098,-0.0005352,0.0080509,-0.0126626,-0.0005172,0.0059194,-0.0014823,0.0033686,-0.0030764,-0.0066882,-0.0025225,0.0037343,0.0012171,-0.0012694
Montenegrin,0.1263436,0.136487,0.039673,0.013889,0.0318826,0.0074744,0.004512,0.0058612,0.0001228,0.002515,-0.001429,-0.0006592,-0.000684,0.0105144,-0.0129208,0.0029436,0.0087096,-0.0013428,0.0070894,-0.0012758,-0.0048414,0.0012364,0.005497,0.0016388,0.0007904
Albanian,0.1181863,0.1417511,0.0155562,-0.0158808,0.0251586,-0.0074138,0.0033683,0.0018077,0.0006648,0.0161279,0.0011097,0.0015486,-0.0070986,0.0066059,-0.019295,-0.0041544,0.0081273,0.0014992,0.0082962,-0.0059402,-0.0055629,0.0007212,0.0024238,0.0021991,-0.0025248
Bulgarian,0.12475,0.1348622,0.0256442,0.0053618,0.02462,-0.0008368,0.005499,0.0030458,-0.0002862,0.0069978,-0.0033452,0.0008992,0.002111,0.0086426,-0.0143046,-0.0022274,0.0070928,0.0020016,0.0031424,-0.0037018,-0.0046168,-0.000544,0.00281,-0.0024824,-0.0010776
Macedonian,0.124598467,0.143257400,0.023406600,-0.006072400,0.025584200,-0.001059867,0.003525000,0.001184467,-0.000191000,0.010897667,0.001245067,0.001198867,-0.0016254,0.009743733,-0.0153364,-0.001706067,0.006684333,-0.000726333,0.006695533,-0.0065114,-0.009782733,0.000404,0.003804267,0.000417733,-0.000478933
Greek_Peloponnese,0.117393,0.1445745,0.0071996,-0.0279395,0.0195001,-0.0102429,0.003429,8.4e-05,0.0019895,0.0140653,0.0040523,0.000872,-0.0016691,0.0049669,-0.0158114,0.0002169,0.0066851,0.0010192,0.0063877,-0.0044055,-0.0066928,0.0010004,0.003815,0.0010133,-0.0018506
Greek_Thessaly,0.119135,0.1452207,0.013702,-0.020026,0.0188753,-0.0051133,0.0005483,0.000769,0,0.01877,0.0026523,0.0040963,-0.0072843,0.0056883,-0.0090933,-0.0041987,0.0021297,-0.000549,0.0041483,-0.005336,-0.0069877,0.0012777,0.0035743,0.0014057,-0.003353
Greek_Kos,0.1076261,0.1464618,-0.0220407,-0.0538692,0.0041031,-0.018252,0.0008617,-0.0036921,-0.0059994,0.0167861,0.0026524,0.0031472,-0.0039148,0.0009634,-0.0106012,0.0029906,0.0100107,0.0009432,0.0037151,-0.0036407,-0.0038128,0.0016762,-0.0001778,-0.0014059,-0.0021157
Greek_Crete,0.1079839,0.145505,-0.0145229,-0.0451942,0.0067765,-0.015247,0.0022748,-0.0018484,-0.002579,0.0128804,0.0019698,0.0015826,-0.0017602,0.0039746,-0.0115539,-0.0023018,0.0046704,0.0009527,0.0038451,-0.0048911,-0.0044534,0.0009002,0.0006446,0.0014423,-0.001692


Portuguese,0.1061741,0.1446114,0.0360075,-0.0044316,0.042137,-0.0040494,-0.0037412,0.0027598,0.0255982,0.029741,-0.000734,0.0074214,-0.0127373,-0.0112631,0.0125568,-0.0003978,-0.0011316,-0.0008869,-0.005762,-0.0002452,0.0011379,-0.002572,0.0013015,-0.0011326,-0.0001053
Basque_French,0.128051,0.1520246,0.0551727,0.0128231,0.0564104,0.0025099,-0.00141,0.0033229,0.0305968,0.0412036,-0.0092724,0.0103408,-0.0209761,-0.0140237,0.0132192,-0.0019359,-0.0119173,0.0030278,-0.0007165,-0.0044144,0.010157,0.0024855,-0.0086397,-0.0091698,7.18e-05
Basque_Spanish,0.1319083,0.1504111,0.0582862,0.0090081,0.0575834,0.0001239,-0.0033162,0.0004102,0.0313603,0.0455997,-0.0045288,0.0116063,-0.0227781,-0.0187627,0.0179452,0.0037271,-0.0058094,0.0027731,-0.0018156,-6.96e-05,0.0087901,0.0024868,-0.0068883,-0.0090371,0.0022619
Spanish_Valencia,0.1179965,0.1446283,0.0408548,-0.0008882,0.0464702,-0.0014408,-0.0019975,0.0016346,0.0231624,0.0332581,-0.0030042,0.0083301,-0.0128593,-0.0123401,0.0096022,-0.0008507,-0.0046938,-0.0017209,-0.001236,-0.0009171,0.0055735,-0.0013706,-0.0020438,-0.0051411,0.0020557
French_Corsica,0.1203273,0.1518216,0.0195294,-0.0230022,0.0350394,-0.0100401,-0.0018464,-0.0005604,0.0157044,0.0337918,-0.0006031,0.0074612,-0.0132308,-0.0071074,-0.004537,-0.0005968,0.0040419,0.0012489,0.0024511,-0.0030461,-0.0003923,0.0002474,-0.0025793,-0.0013858,-0.0005303
French_Paris,0.1271714,0.1414355,0.0504313,0.0239901,0.0409027,0.0093048,0.0027985,0.0054544,0.0116022,0.0181739,-0.0077061,0.0068121,-0.0129065,-0.0103844,0.0137695,0.0060631,-0.0018255,0.0046644,0.0027082,-0.0009778,0.0022007,0.0035747,-0.0012325,0.0111405,-0.0019051
French_South,0.1261811,0.1447856,0.0496183,0.0088594,0.0506907,0.0026694,0.0002686,0.0022747,0.0276401,0.0386601,-0.0067741,0.0073649,-0.021853,-0.0135659,0.0103729,0.0027653,-0.0051034,0.0017917,0.0002154,0.0019116,0.0049911,-0.0008301,-0.0052646,-0.0075569,-0.00142
Swiss_French,0.1273395,0.1448398,0.0414364,0.0102149,0.0411615,0.0064842,0.0008224,0.0028845,0.0118111,0.0194308,-0.000751,0.0065942,-0.0159622,-0.0125238,0.0083468,0.0021712,0.0057533,-0.0016154,0.003661,-0.0003439,0.0047886,0.0036478,0.0016638,0.0025756,-0.0004791
Swiss_German,0.127624,0.1407779,0.0460088,0.0227311,0.0406229,0.0073906,0.0027906,0.0051344,0.0086409,0.0121641,-0.0015224,0.005264,-0.0149402,-0.008808,0.0083638,-0.0011269,-0.0025912,0.0012192,0.0053735,-0.0009222,0.0019341,0.0035086,-0.0025574,0.006974,-0.0025596
Swiss_Italian,0.1221703,0.1411583,0.0232557,-0.014535,0.026774,-0.0027887,0.000235,0.0036923,0.00934,0.0202283,0.0007577,0.003097,-0.0111993,-0.0034867,-0.0030763,0.0019447,0.0061283,-0.0024913,0.002095,-0.0022927,0.002329,-0.0002473,-0.001479,0.0061453,-0.0005187
Italian_Lazio,0.1146768,0.1525832,0.006694,-0.0350455,0.0213115,-0.0115738,0.0024675,0.000519,0.0068512,0.0234627,0.001299,0.0028475,-0.007656,-0.0070875,-0.0068538,0.0041432,-0.000228,-0.0005068,0.0018225,-0.005065,-0.002745,0.0033695,-0.0003697,-0.0003613,-0.000479
Italian_Lombardy,0.1245224,0.147861,0.0279822,-0.0137598,0.0327446,-0.007753,0.000658,0,0.0103898,0.0299232,-0.0025008,0.008932,-0.0143606,-0.0047066,-0.0008686,-0.0078758,-0.0083448,0.0017738,0.0034442,-0.004102,-0.002246,0.0012366,-0.0006656,0.0030602,0.0008382
Sardinian,0.1216872,0.1672855,0.0284897,-0.0506523,0.060151,-0.0221337,-0.0039523,0.0024964,0.0415743,0.077351,-5.9e-05,0.0166487,-0.0286645,-0.012974,-0.013572,-0.0030013,0.0114027,-0.0013475,0.0016798,-0.0129947,-0.0021212,-0.0011018,-0.010084,-0.021383,0.0003375